Operational definition

Ecological pool category.

Organic matter containing chemical free energy that remains within an ecosystem pool for an ecologically significant residence time.

Examples include:

Unlike behavioral storage, ecosystem-scale storage need not be intentional or retrievable by the organism that produced it. Glycogen and starch are widespread carbon and energy reserves; lipids and microbial polymers provide other storage forms, while storage theory treats reserve accumulation as an allocation strategy under future uncertainty. (PubMed Central (PMC))