108–115 — OUTPUTS OF A LIVING ENERGY SYSTEM
The Outputs of a Living Energy System layer identifies the matter and energy that remain as newly formed stocks within the focal system or leave that system as a consequence of metabolism, growth, interaction, reproduction, waste removal, and dissipation.
Operational definition
Layer type: Boundary-output and retained-stock layer Governs: Items 108–115
Governing question
What physically remains or crosses the boundary after acquisition, conversion, assimilation, and allocation?
Included outputs
- retained biomass;
- retained reserves;
- reproductive export;
- partner export;
- secreted products;
- excreted or egested material;
- terminal metabolic products;
- heat.
Stock-versus-output qualification
Not every child is an outward flow.
- Retained biomass is an internal stock increase.
- Retained reserves are an internal stock increase.
- Reproductive export is an outward flow.
- Heat is an outward energy transfer.
The layer therefore records system outcomes, including both retained products and exported products.
Inclusion rule
A quantity belongs here when it is a physically identifiable endpoint of the focal accounting interval.
The entry should specify:
- material or energy form;
- stock or flow;
- destination;
- chemical composition;
- time interval;
- whether it remains biologically usable.
Function-versus-output distinction
The same process can be described differently across layers.
For example:
- allocation: reproduction;
- life-history event: reproductive export;
- physical output: egg containing a measured quantity of biomass and chemical free energy;
- ecological transfer: offspring or consumer resource.
These are not duplications if they answer different questions and are not summed indiscriminately.
Terminal-product rule
A product is terminal only relative to the focal system and stated conditions.
For one organism:
CO_2
may be a terminal metabolic product.
For an autotroph, the same carbon dioxide may be an input.
Heat rule
Heat represents energy transfer in a dispersed thermal form. It is not the destruction of energy, but it ordinarily represents a reduction in the remaining capacity to drive organized biological work under ambient conditions.
108. Retained biomass
- new cells
- new tissue
- structural material
- reproductive structures
109. Retained reserves
- carbohydrate
- lipid
- storage polymers
- stored reduced compounds
110. Reproductive export
- offspring
- eggs
- seeds
- spores
- propagules
- parental provisioning
111. Partner export
- exchanged metabolites
- leaked substrates
- symbiotic products
- colony-level transfers
112. Secreted products
- enzymes
- toxins
- mucus
- extracellular matrix
- organic acids
- gases
- signaling compounds
113. Excreted or egested material
- feces
- nitrogenous waste
- mineral waste
- undigested matter
- dead cells
- shed structures
114. Terminal metabolic products
Depending upon the metabolism:
- carbon dioxide
- water
- methane
- hydrogen sulfide
- nitrogen gas
- nitrous oxide
- organic acids
- alcohols
- molecular hydrogen
- reduced metals
- oxidized minerals
115. Heat
Every real transformation produces entropy and ultimately degrades some capacity to perform useful work.
The precise statement is:
Energy is conserved; usable free energy is progressively degraded.